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First of all, MAC OS server requires a static IP number to work fine.
Secondly if the server is behind a router (and it must be if your INTERNET IP number is not static), than forward the incoming mail through the router to the server.
If you want to access the mail from the internet too, forward all related roouter ports to the server also.
You do not need to have an active DNS server (in the mac os server) to use the mail functionality of the server. Having an DNS server makes access easier, but is not a requirement at all.
I use the same setup as describe above (thus a semi-static internet IP-number for incoming mail) and a router with port forwarding to the MAIL server.
You can check the proper working of the mail server behind the router by sending emails directly to it using its static ip-number. (outgoing mail = ip-number e.g. 192.168.1.2). The server must not get its ip-number from the DHCP server in your router, but it must be static (and outside the range of numbers issued by the DHCP server of your router).
Good luck, Kees
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